Predictions & Data for this entry
| Model: hep | climate: Cfb, Dfb, Dfc | migrate: | phylum: |
| COMPLETE = 4.0 | ecozone: TH | food: beH, beD, beS | class: |
| MRE = 0.148 | habitat: 0eFl, 0eFp, 0eFm, eiTg | gender: D | order: |
| SMSE = 0.082 | embryo: Fv | reprod: Os | family: |
Zero-variate data
| Data | Observed | Predicted | (RE) | Unit | Description | Reference |
|---|---|---|---|---|---|---|
| ab | 10 | 8.327 | (0.1673) | d | age at birth | Wiki |
| tj_26 | 23 | 25.9 | (0.1262) | d | time since birth at emergence at 26 C | gaia2019 |
| tj_22 | 30 | 35.48 | (0.1827) | d | time since birth at emergence at 22 C | gaia2019 |
| tj_18 | 47 | 50.72 | (0.0792) | d | time since birth at emergence at 18 C | gaia2019 |
| tj_13 | 142 | 122.6 | (0.1367) | d | time since birth at emergence at 13 C | gaia2019 |
| am | 21 | 21.14 | (0.00672) | d | life span as (sub)imago | Wiki |
| L0 | 0.0167 | 0.01657 | (0.008041) | cm | egg length | Degr1960 |
| Lb | 0.0554 | 0.06618 | (0.1945) | cm | length at hatching | Degr1960 |
| Lj | 0.93 | 0.9154 | (0.01569) | cm | body length female imago | Cian1979 |
| Lj_m | 0.84 | 0.8614 | (0.02544) | cm | body length male imago | Cian1979 |
| Lj_w | 1.03 | 0.6606 | (0.3586) | cm | wing length female imago | Swee2018 |
| Wd0 | 0.0006 | 0.0008024 | (0.3373) | mg | initial dry weight | guess |
| Wdb | 0.0009 | 0.0006701 | (0.2554) | mg | dry weight at birth | Swee2018 |
| Wwj | 13.6 | 11.13 | (0.1813) | mg | wet weight female subimago | Cian1979 |
| Wwj_m | 8.2 | 9.311 | (0.1355) | mg | wet weight male subimago | Cian1979 |
| L1 | 0.1962 | 0.06618 | (0.6627) | cm | body length of instar 1 | gaia2019 |
| L2 | 0.263 | 0.294 | (0.118) | cm | body length of instar 2 | gaia2019 |
| L3 | 0.3654 | 0.3732 | (0.02132) | cm | body length of instar 3 | gaia2019 |
| L4 | 0.4988 | 0.5105 | (0.02338) | cm | body length of instar 4 | gaia2019 |
| L5 | 0.6015 | 0.7123 | (0.1842) | cm | body length of instar 5 | gaia2019 |
| L6 | 0.6845 | 0.7602 | (0.1105) | cm | body length of instar 6 | gaia2019 |
| L7 | 0.6703 | 0.7655 | (0.142) | cm | body length of instar 7 | gaia2019 |
| L8 | 0.6745 | 0.765 | (0.1341) | cm | body length of Nymph instar 1 | gaia2019 |
| L9 | 0.6695 | 0.7274 | (0.0865) | cm | body length of Nymph instar 2 | gaia2019 |
| Wd1 | 0.02078 | 0.0006716 | (0.9677) | mg | dry weight of instar 1 | gaia2019 |
| Wd2 | 0.06111 | 0.0589 | (0.03615) | mg | dry weight of instar 2 | gaia2019 |
| Wd3 | 0.1175 | 0.1204 | (0.02466) | mg | dry weight of instar 3 | gaia2019 |
| Wd4 | 0.3216 | 0.3082 | (0.04172) | mg | dry weight of instar 4 | gaia2019 |
| Wd5 | 0.8554 | 0.8374 | (0.02103) | mg | dry weight of instar 5 | gaia2019 |
| Wd6 | 1.091 | 1.018 | (0.06741) | mg | dry weight of instar 6 | gaia2019 |
| Wd7 | 1.105 | 1.039 | (0.05931) | mg | dry weight of instar 7 | gaia2019 |
| Wd8 | 1.024 | 1.037 | (0.01255) | mg | dry weight of Nymph instar 1 | gaia2019 |
| Wd9 | 0.9199 | 0.8919 | (0.03047) | mg | dry weight of Nymph instar 2 | gaia2019 |
| Lh1 | 0.033 | 0.009632 | (0.7081) | cm | head length of instar 1 | gaia2019 |
| Lh2 | 0.0422 | 0.0428 | (0.01414) | cm | head length of instar 2 | gaia2019 |
| Lh3 | 0.0584 | 0.05432 | (0.06987) | cm | head length of instar 3 | gaia2019 |
| Lh4 | 0.0732 | 0.0743 | (0.01504) | cm | head length of instar 4 | gaia2019 |
| Lh5 | 0.0992 | 0.1037 | (0.04512) | cm | head length of instar 5 | gaia2019 |
| Lh6 | 0.1092 | 0.1106 | (0.01325) | cm | head length of instar 6 | gaia2019 |
| Lh7 | 0.1153 | 0.1114 | (0.03367) | cm | head length of instar 7 | gaia2019 |
| Lh8 | 0.1092 | 0.1113 | (0.01965) | cm | head length of Nymph instar 1 | gaia2019 |
| Lh9 | 0.1153 | 0.1059 | (0.08172) | cm | head length of Nymph instar 2 | gaia2019 |
Uni- and bivariate data
| Data | Figure | Independent variable | Dependent variable | (RE) | Reference |
|---|---|---|---|---|---|
| tL_f |   | time | length | (0.2018) | Cian1979 |
| tL_m | | time | length | (0.191) | Cian1979 |
| tW_f |   | time | wet weight^{1/3} | (0.2262) | Cian1979 |
| tW_m | | time | wet weight^{1/3} | (0.154) | Cian1979 |
| WJO_f |   | wet weight | O_2 consumption | (0.2908) | Cian1979 |
| WJO_m | | wet weight | O_2 consumption | (0.3042) | Cian1979 |
| tL_f1 |   | time since start of experiment | length | (0.1317) | gaia2016 |
| tL_m1 | | time since start of experiment | length | (0.1295) | gaia2016 |
| tL_c |   | time | length | (0.04754) | gaia2016 |
| tL_w | | time | length | (0.04991) | gaia2016 |
| tL_h1 |   | time since start of experiment | head length | (0.08156) | gaia2019 |
| tL_h2 | | time since start of experiment | head length | (0.06943) | gaia2019 |
| tL_h3 | | time since start of experiment | head length | (0.09542) | gaia2019 |
| tL_h4 | | time since start of experiment | head length | (0.3386) | gaia2019 |
| tL_h5 | | time since start of experiment | head length | (0.09942) | gaia2019 |
| tS |   | starvation time | survival | (0.1205) | Wage2015 |
| tS_2 | | starvation time | survival | (0.05418) | Wage2015 |
| tS_3 | | starvation time | survival | (0.03041) | Wage2015 |
| Lw_N |  | length female imago | No. of hatchlings | (0.2776) | Swee2018 |
| Lw_Wd |  | length female imago | dry weight | (0.1599) | Swee2018 |
| Ttj |  | Temperature | time since birth at emergence | (0.3811) | Swee2018 |
| Ttj_g |  | Temperature | time since birth at emergence | (0.1672) | gaia2019 |
| tL |  | time since birth | length | (0.17) | GuptMich1993 |
| tL_g |  | time | length | (0.03129) | gaia2016 |
Pseudo-data at Tref = 20°C
| Data | Generalised animal | Cloeon dipterum | Unit | Description |
|---|---|---|---|---|
| v | 0.02 | 0.00299 | cm/d | energy conductance |
| p_M | 18 | 809 | J/d.cm^3 | vol-spec som maint |
| k_J | 0.002 | 0.002 | 1/d | maturity maint rate coefficient |
| k | 0.3 | 0.01105 | - | maintenance ratio |
| kap | 0.8 | 0.5975 | - | allocation fraction to soma |
| kap_G | 0.8 | 0.7957 | - | growth efficiency |
| kap_R | 0.95 | 0.95 | - | reproduction efficiency |
Discussion
- Males are assumed to differ from females by {p_Am} only
- tL and tW data are from wild-catch of unknown time since birth
- Food (periphyton) in experiments of gaia2016 was of high quality and therefore f = 1 was assumed
- hep-model is not entirely able to represent the sigmoidal growth of Cloeon larvae: late instars larvae do not grow in lenght anymore probably due to decreased feeding
- At low food, larval development time is prolonged, and size at emergence seems rather conserved across food levels
- Dry weight over temperature (Swee2018 data) cannot be predicted by the model
- mod_2: Many new data sets added
Facts
- The only ovoviviparous ephemeropteran species in Europe; Females lay eggs 10-14 days after mating, and the eggs hatch as soon as they hit the water (Ref: Wiki)
- Dry weight in mg = 0.001 * (body length in mm)^3.6802 (Ref: Cian1979)
- Females are 10% longer then males after larval stage 7 (Ref: Cian1979)
- Weights-at-stages and moulting frequency are unaffected by food quality, but duration is (so is the total number of moults) (Ref: Cian1979)
- Larval moults are 5% of dry body mass, 7.5% (female) or 11.5% (male) in N2, 13% (f) or 16% (m) in subimago (Ref: Cian1979)
- Wet/dry weight ratio is about 3 (Ref: Cian1979)
Bibliography